| Preface | p. xiii |
| Prologue | p. 1 |
| Morphology, evolutionary history and recent distribution | p. 6 |
| Introduction | p. 6 |
| Morphology | p. 6 |
| Evolutionary origins and relationships | p. 16 |
| Paleontological diversity | p. 20 |
| Distribution of extant species | p. 21 |
| Food and other habitat resources | p. 30 |
| Introduction | p. 30 |
| Food | p. 30 |
| Water and other habitat needs | p. 45 |
| Comparisons with smaller ungulates | p. 50 |
| Space--time patterns of habitat use | p. 53 |
| Introduction | p. 53 |
| Temporal patterning of activities | p. 53 |
| Utilization of space | p. 61 |
| Comparisons with smaller ungulates | p. 67 |
| Body size and nutritional physiology | p. 69 |
| Introduction | p. 69 |
| Metabolic requirements | p. 70 |
| Gut anatomy | p. 71 |
| Food intake and digestion | p. 72 |
| Body size and feeding ecology | p. 82 |
| Introduction | p. 82 |
| Diet quality | p. 82 |
| Foraging time | p. 87 |
| Home range extent | p. 95 |
| Trophic ecology of megaherbivores: summary | p. 98 |
| Social organization and behavior | p. 101 |
| Introduction | p. 101 |
| Group structure | p. 101 |
| Male dominance relations | p. 109 |
| Courtship and mating | p. 116 |
| Responses to predators | p. 124 |
| Comparisons with smaller ungulates | p. 131 |
| Life history | p. 133 |
| Introduction | p. 133 |
| Infancy and juvenilehood | p. 133 |
| Adolescence and puberty | p. 138 |
| Reproduction by females | p. 144 |
| Reproduction by males | p. 151 |
| Mortality and lifespan | p. 152 |
| Comparisons with smaller ungulates | p. 159 |
| Body size and sociobiology | p. 160 |
| Introduction | p. 160 |
| Grouping patterns | p. 160 |
| Male dominance systems | p. 167 |
| Female mate choice | p. 177 |
| Summary | p. 179 |
| Body size and reproductive patterns | p. 181 |
| Introduction | p. 181 |
| Seasonality of reproduction | p. 183 |
| Age at first conception | p. 185 |
| Birth intervals | p. 186 |
| Maternal investment in reproduction | p. 190 |
| Offspring sex ratio | p. 195 |
| Summary | p. 198 |
| Demography | p. 200 |
| Introduction | p. 200 |
| Population structure | p. 200 |
| Population growth | p. 212 |
| Population density and biomass | p. 221 |
| Comparisons with smaller ungulates | p. 225 |
| Community interactions | p. 226 |
| Introduction | p. 226 |
| Impact on vegetation | p. 226 |
| Effects on other large herbivores | p. 239 |
| Comparisons with smaller ungulates | p. 245 |
| Body size and population regulation | p. 246 |
| Introduction | p. 246 |
| Demographic models | p. 248 |
| Interactions with vegetation | p. 257 |
| Dispersal | p. 260 |
| Summary | p. 264 |
| Body size and ecosystem processes | p. 265 |
| Introduction | p. 265 |
| Biomass levels | p. 265 |
| Energy flux | p. 274 |
| Nutrient cycling | p. 277 |
| Ecosystem stability and disturbance | p. 278 |
| Late Pleistocene extinctions | p. 280 |
| Introduction | p. 280 |
| Pattern of extinctions | p. 281 |
| Climatic change | p. 284 |
| Human predation | p. 289 |
| The role of megaherbivores | p. 292 |
| Summary | p. 296 |
| Conservation | p. 297 |
| Introduction | p. 297 |
| Conservation objectives | p. 298 |
| Problems of overabundance | p. 299 |
| Problems of overexploitation | p. 307 |
| Summary | p. 308 |
| Epilogue: the megaherbivore syndrome | p. 309 |
| Faunal patterns | p. 309 |
| Social and life history patterns | p. 312 |
| Demographic patterns | p. 313 |
| Community and ecosystem patterns | p. 314 |
| Appendixes | p. 317 |
| Appendix I | p. 317 |
| Appendix II | p. 325 |
| References | p. 331 |
| Index | p. 364 |
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