| Preface | p. xi |
| Introduction to the Fernow Watershed Acidification Study | |
| Introduction | p. 1 |
| Atmospheric Deposition at Fernow | p. 2 |
| Theory of Acidification by Atmospheric Deposition | p. 3 |
| Fernow Research Themes | p. 11 |
| Nutrient Cation Mobilization | p. 11 |
| Cation Depletion/Aluminum Mobilization | p. 12 |
| Biomass Growth Cycles | p. 13 |
| Steady States | p. 14 |
| Book Organization | p. 14 |
| References | p. 15 |
| Fernow and the Appalachian Hardwood Region | |
| Regional Description | p. 17 |
| Disturbance Agents | p. 18 |
| Vegetation Description | p. 19 |
| Early Settlement and Industrial Development | p. 20 |
| Early Fernow History | p. 20 |
| Fernow Research | p. 22 |
| Fernow Geology and Soils | p. 24 |
| Fernow Forest Composition | p. 26 |
| Fernow Climate | p. 28 |
| Study Watersheds | p. 28 |
| Water Instrumentation | p. 32 |
| Watershed Acidification Treatments | p. 33 |
| References | p. 34 |
| Soil Chemical Response to Experimental Acidification Treatments | |
| Introduction | p. 41 |
| Methods | p. 42 |
| Comparisons between WS3 and WS4 | p. 42 |
| Comparisons between WS3 and WS7 | p. 44 |
| Effects on Soils on WS9 | p. 46 |
| Results | p. 47 |
| Pretreatment Soil Characteristics - WS3, WS4, WS7 | p. 47 |
| Pretreatment Soil Characteristics - WS9 | p. 49 |
| Treatment Effects | p. 49 |
| Comparisons between WS3 and WS4 | p. 49 |
| Comparisons between WS3 and WS7 | p. 56 |
| Treatment Effects WS9 | p. 58 |
| Soil N Cycling | p. 62 |
| Discussion | p. 63 |
| Conclusion | p. 65 |
| References | p. 66 |
| Soil Water and Stream Water Chemical Responses | |
| Introduction | p. 71 |
| Methods | p. 72 |
| Soil Water Sampling | p. 72 |
| Stream Water Sampling | p. 72 |
| Chemical Analyses | p. 76 |
| Data Analyses | p. 77 |
| Soil Water | p. 79 |
| WS4 and WS3 Responses | p. 79 |
| WS9 Responses | p. 96 |
| Stream Water | p. 103 |
| Concentrations in Baseflow | p. 105 |
| WS4 Responses | p. 105 |
| WS3 Responses | p. 108 |
| WS9 Responses | p. 112 |
| Concentrations at Peakflow | p. 118 |
| WS4 Responses | p. 118 |
| WS3 Responses | p. 118 |
| WS9 Responses | p. 121 |
| Discussion | p. 124 |
| Application of Base Cation Results | p. 124 |
| Application of Nitrate Responses | p. 126 |
| Regional Susceptibility to Acidification | p. 127 |
| Summary and Conclusion | p. 129 |
| References | p. 131 |
| Vegetation and Acidification | |
| Introduction | p. 137 |
| Vegetation Description | p. 139 |
| Fernow | p. 139 |
| Stand Conditions - WS3 and WS7 | p. 139 |
| Herbaceous Layer Characteristics - WS3 and WS7 | p. 140 |
| Stand Conditions - WS9 | p. 142 |
| Methods | p. 143 |
| Tree and Stand Growth | p. 143 |
| Growth Plot Analyses | p. 143 |
| Tree Core Analysis | p. 144 |
| Foliar Chemistry | p. 146 |
| Dendrochemistry | p. 146 |
| Herb Layer Dynamics | p. 147 |
| Results | p. 148 |
| WS3 vs. WS7 (Fernow) Results | p. 148 |
| Tree Growth | p. 148 |
| Growth Plots | p. 148 |
| Tree Cores | p. 150 |
| Overstory Foliar Chemistry | p. 156 |
| Dendrochemistry | p. 159 |
| Herb Layer Effects | p. 165 |
| WS9 (Clover Run) Results | p. 171 |
| Tree Growth | p. 171 |
| Foliar Chemistry | p. 173 |
| Dendrochemistry | p. 173 |
| Discussion | p. 174 |
| Conclusions | p. 182 |
| References | p. 184 |
| Response of Salamanders to Experimental Acidification Treatments | |
| Introduction | p. 189 |
| Salamanders in the Fernow Experimental Forest | p. 190 |
| Biology and Ecology of Salamanders | p. 190 |
| Effects of Disturbance on Salamanders | p. 191 |
| Salamander Research on the Fernow Experimental Forest | p. 193 |
| Methods | p. 194 |
| Effects of Watershed Acidification | p. 194 |
| Effects of Clearcutting | p. 195 |
| Results | p. 195 |
| Effects of Watershed Acidification on Salamanders | p. 195 |
| Effects of Clearcutting on Salamanders | p. 197 |
| Discussion | p. 198 |
| Conclusions | p. 199 |
| References | p. 200 |
| Acidification and Nutrient Cycling | |
| Introduction | p. 207 |
| Nitrogen Cycling | p. 208 |
| Inputs | p. 208 |
| Outputs | p. 210 |
| Budgets | p. 211 |
| Internal Cycling | p. 212 |
| Sulfur Cycling | p. 216 |
| Inputs | p. 216 |
| Outputs | p. 217 |
| Budgets | p. 219 |
| Internal Cycling | p. 220 |
| Calcium Cycling | p. 221 |
| Inputs | p. 221 |
| Outputs | p. 221 |
| Budgets | p. 221 |
| Internal Cycling | p. 223 |
| Magnesium Cycling | p. 226 |
| Inputs | p. 226 |
| Outputs | p. 226 |
| Budgets | p. 226 |
| Internal Cycling | p. 226 |
| Potassium Cycling | p. 228 |
| Inputs | p. 228 |
| Outputs | p. 228 |
| Budgets | p. 228 |
| Internal Cycling | p. 228 |
| Conclusions | p. 229 |
| References | p. 232 |
| Implications for Forest Ecosystem Sustainability and Resiliency | |
| Introduction | p. 237 |
| Methods | p. 238 |
| Model Description: Focus on N Deposition | p. 238 |
| Nitrogen Deposition Scenarios | p. 239 |
| Results | p. 243 |
| N Deposition Modeling and Mapping | p. 243 |
| Net Primary Productivity, NPP | p. 245 |
| Water Yield | p. 245 |
| Current Forest N Leaching Losses and Retention | p. 245 |
| Comparison with Field Data | p. 252 |
| Conclusions | p. 253 |
| References | p. 254 |
| Conclusions and Recommendations from the Fernow Watershed Acidification Study | |
| Nutrient Cation Mobilization | p. 259 |
| Cation Depletion/Aluminum Mobilization | p. 261 |
| Biomass Growth Cycles | p. 263 |
| Steady States | p. 264 |
| Implications for Forest Sustainability and Resiliency | p. 265 |
| Research Recommendations | p. 266 |
| References | p. 268 |
| Appendices | p. 269 |
| Index | p. 277 |
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