| Preface | p. xiii |
| Patterns of culture? | p. 1 |
| Introduction | p. 1 |
| The prey | p. 2 |
| Six key sites | p. 3 |
| Other sites | p. 6 |
| Non-human culture? | p. 10 |
| Studying chimpanzees | p. 15 |
| Introduction | p. 15 |
| Development of chimpanzee research | p. 16 |
| Studies in nature | p. 16 |
| Studies in capitivity | p. 20 |
| Sites of study | p. 23 |
| Eastern chimpanzees | p. 23 |
| Central-western chimpanzees | p. 25 |
| Western chimpanzees | p. 27 |
| Captive chimpanzees | p. 28 |
| Methods of study | p. 29 |
| Studies in nature | p. 30 |
| Methodological issues | p. 34 |
| Studies in capitivity | p. 36 |
| Collecting data | p. 38 |
| Conclusion | p. 39 |
| Chimpanzees as apes | p. 40 |
| Introduction | p. 40 |
| Sources and methods | p. 41 |
| Patterns of tool-use | p. 44 |
| Chimpanzee | p. 44 |
| Bonobo | p. 47 |
| Orang-utan | p. 49 |
| Highland gorilla | p. 51 |
| Lowland gorilla | p. 51 |
| Gibbon | p. 52 |
| Socio-ecology | p. 53 |
| Brain | p. 55 |
| Hands | p. 57 |
| Mind | p. 57 |
| Apes and their tools | p. 59 |
| Ancestral hominoids | p. 62 |
| Cultured chimpanzees? | p. 65 |
| Introduction | p. 65 |
| Gombe and Kasoje compared | p. 65 |
| Case study: Grooming | p. 67 |
| Defining culture | p. 72 |
| Japanese macaques | p. 77 |
| Additional conditions for culture | p. 78 |
| Chimpanzees as culture-bearers? | p. 79 |
| Culture denied? | p. 82 |
| Chimpanzee sexes | p. 88 |
| Introduction | p. 88 |
| Sex or gender? An aside | p. 89 |
| Sex differences in diet: invertebrates | p. 89 |
| Case study: Termite-fishing | p. 90 |
| Chimpanzees, tools and termites | p. 92 |
| Case study: Ant-dipping | p. 93 |
| Chimpanzees and ants | p. 97 |
| Sex differences in diet: meat | p. 99 |
| Case study: Mammals as prey | p. 99 |
| Carnivory elsewhere | p. 102 |
| Sex and faunivory | p. 103 |
| Nut-cracking | p. 105 |
| Food-sharing | p. 106 |
| Case-study: Banana-sharing | p. 107 |
| Other food sharing | p. 111 |
| Other apes | p. 112 |
| Origins of sexual division of labour | p. 113 |
| Origins of tool-use | p. 115 |
| Chimpanzees and foragers | p. 121 |
| Cautionary note | p. 121 |
| Why compare chimpanzees and hunter-gatherers? | p. 121 |
| Ideal versus actual comparisons | p. 123 |
| Hot, dry and open habitats: humans and apes compared | p. 127 |
| Case study: Tasmanian humans and Tanzanian apes | p. 131 |
| Oswalt's taxonomy | p. 131 |
| Choosing samples | p. 134 |
| Tasmanian aborigines | p. 136 |
| Tanzanian chimpanzees | p. 139 |
| Subsistants compared | p. 141 |
| Similarities and differences | p. 144 |
| Diet | p. 144 |
| Food acquisition and processing | p. 146 |
| Conclusions | p. 149 |
| Chimpanzees compared | p. 150 |
| Introduction | p. 150 |
| Difficulties of comparison: eating meat | p. 150 |
| Chimpanzee insectivory | p. 153 |
| Termites | p. 155 |
| Ants | p. 159 |
| Honey | p. 162 |
| Explaining variation | p. 166 |
| Case study: 'Fishing' for termites | p. 168 |
| Hammers and anvils | p. 173 |
| Chimpanzee ethnology | p. 177 |
| Cataloguing tool-kits | p. 177 |
| Non-subsistence technology | p. 181 |
| Regional and local patterns | p. 189 |
| Innovation | p. 193 |
| The invention of termite-fishing | p. 195 |
| Cross-cultural chimpanzees? | p. 196 |
| Chimpanzees as models | p. 198 |
| Kinds of models | p. 198 |
| Models of what? | p. 200 |
| Stone artefacts | p. 202 |
| Why have palaeo-anthropologists ignored other primates? | p. 207 |
| Another cautionary note | p. 212 |
| Conclusions | p. 214 |
| What chimpanzees are, are not, and might be | p. 215 |
| Introduction | p. 215 |
| Conceiving of chimpanzees | p. 215 |
| Evolutionarily relevant gaps | p. 217 |
| Hunting | p. 218 |
| Gathering | p. 218 |
| Food processing | p. 219 |
| Communication | p. 222 |
| Unanswered questions | p. 223 |
| Conclusions | p. 230 |
| Scientific names | p. 231 |
| References | p. 233 |
| Author index | p. 264 |
| Subject index | p. 270 |
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